| 1973 |
Dipl. Biochem., University of Tübingen |
| 1976 |
Dr. rer. nat., Max Planck Institute for Biochemistry, Martinsried |
| 1977-1978 |
Postdoctoral fellow at the Biocenter Basel |
| 1978-1979 |
Research assistant at the University of Heidelberg |
| 1980-1991 |
Research fellow at the Max Planck Institute for Psychiatry, Martinsried |
| 1985/1992 |
Habilitation (Zoology/Apl. Prof.), Ludwig-Maximilians-University Munich |
| 1995 |
Apl. Prof., J.-W.-Goethe University Frankfurt |
| since 1991 |
Research group leader at MPIH |