| 1968 |
Dipl. Phys. (Physiological Optics), LM University of München |
| 1972 |
Dr. rer. nat. (Neurobiology), LM University and Max Planck Institute for Psychiatry, München |
| 1972 |
Postdoc, Cell Biophysics Unit of the Medical Research Council (MRC), London |
| 1973 |
Dept. Physiol. J.C.S.M.R. Australian National University, Canberra |
| 1978 |
Habilitation, University of Konstanz |
| 1977 |
Groupleader, Friedrich-Miescher-Labor/Tübingen |
| since 1981 |
Director, Department of Neuroanatomy |
| since 1982 |
Honorary Professor, University of Mainz |
| since 11/2008 |
Director Emeritus |